Jonathan Lindsay Penner is an American actor, writer and film producer known for starring in The Last Supper and the television series Rude Awakening and The Naked Truth, as well as for appearing on the television show Survivor. Penner was born in New York City. He is related to actor Jason Alexander, who is married to Daena Title - the cousin of Penner's wife, Stacy Title. Penner has starred on three television series: Grapevine, The Naked Truth and Rude Awakening and guest-starred in Seinfeld, CSI, CSI: NY, The Tick, Melrose Place, LateLine, Arrested Development, and The Nanny, where he... appeared in several episodes as Danny Imperiali, whose firing and dumping of fiance Fran Fine kicked off the entire series. Penner has appeared in many films, with featured roles in The Last Supper, Anarchy TV and Down Periscope. He was nominated for an Academy Award in 1994 for the short film Down on the Waterfront, which he co-produced, co-wrote, and starred in.
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